For Mira Simon (Jessica Szohr), it feels like a prank when all of her dreams start coming true on April Fool's Day. An April Fool, however, she is not. After three of her own tricks materialize, Mira finds herself promoted to the perfect job, dating the perfect man, and living in the perfect home. Her huge promotion even allows her to work directly with her professional hero, CEO Erin Billings (Deidre Hall) and her friend, Jonah (Benjamin Hollingsworth), on a massive social media website relaunch.
Single mother Ryan (Tricia Helfer) has just about given up on dating after her divorce, happily accepting her young son as the most important man in her life. That all changes when Ryan's brother Owen (JT Hodges), also feeling unlucky in love after a bad breakup, swaps his home in their small North Carolina town with New York City adman Sean (Mark Lutz). Ryan feels an instant spark with her new neighbor, but Sean, who signed up for the temporary home swap after a proposal gone wrong, is out of his comfort zone in his new rural surroundings.